The remains of recently identified Greek soldiers who fought in Cyprus against invading Turkish troops nearly half a century ago were returned to their families on Thursday.

Cypriot President Nikos Christodoulides attended a funeral service at Ayios Panteleimonas Orthodox Church in the capital, Nicosia, for the 15 soldiers before their remains were contained in Greek flag-draped coffins.

Mr Christodoulides said it was the least the state could do to honour and pay respects to the memory of those who died.

Eight of the 15 soldiers will be reinterred in Greece.

The families of another six opted to have their remains reinterred in a mass grave in Nicosia which stands as the country’s prime monument for the war.

No family members have been located for one of the soldiers, according to the state broadcaster.

Turkey invaded Cyprus in July 1974, a week after supporters of union with Greece mounted a coup backed by the Greek junta then ruling the country.

The invasion resulted in Cyprus’s ethnic divide, with Turkish Cypriots later declaring independence that is only recognised by Turkey, which still maintains more than 35,000 troops in the breakaway north.

Of the 2,002 people who disappeared in 1974 and the preceding decade amid ethnic violence, the remains of 1,033 have been identified and returned to their families since UN-led search efforts began in earnest in 2006.

UN officials said this marks the second-best success rate in the world, after the former Yugoslavia.

A total of 769 Greek Cypriots and 200 Turkish Cypriots are still listed as missing and officials say the passage of time poses a huge challenge.